First records of some Oribatid mite species (Acari, Oribatida) from Ukraine

: Four species of oribatid mites known mainly from central Europe: Oppiella hygrophila (Mahunka, 1987), Oxyoppia europaea Mahunka, 1982, Achipteria cf. quadridentata Willmann, 1951 and Ceratozetes cf. psammophilus Horak, 2000 are recorded from Ukraine for the first time. The new records of the first three species extend the known areas of their occurrence to the east of Europe (Zakarpattia region).


INTRODUCTION
The contemporary fauna of the oribarid mites of the world consists of more than 11 thousand species (1278 genera and 163 families) (Subias 2018).More than 700 species (ca.7 % of the world fauna are known from Ukraine) (Yaroshenko 2000).But the fauna is being studied, the number of taxa is increasing (Melamud 2003, 2008, 2009, Hushtan 2015, 2017).Transcarpathia (Zakarpattia) is one of the richest in the taxonomic relation of the regions of Ukraine.No less interesting for taxonomists is the Transcarpathian lowland.The oribatid mites remained poorly studied in this territory (Hushtan 2017).While identifying the material from the samples collected in the Transcarpathian plain, four oribatid species, previously known only from neighboring Central European countries, were found.

MATERIAL AND METHODS
All of these studies are conducted according to generally accepted methods of soil zoology (Krantz & Walter 2009, Potapov & Kuznetsova 2011).Samples were taken using a cenometer (125 cm 3 ) (Potapov & Kuznetsova 2011).Extraction of mites from soil was carried out using Tullgren funnel metod.Permanent preparations were prepared.Identification of the specimens was carried out using a microscope (Olympus BX 42) and the key of Weigmann & Miko (2006).In the four types of habitats: hygrophilic, hydromeliorated, mesophilic grasslands and floodplain meadow, were found thirty five specimens belonging to four species of the Oribatida which were previously unknown from Ukraine.I follow the classification provided by Weigmann & Miko (2006).Distribution of species was summarized based on the works of Weigmann &Miko (2006) andSubias (2018).
The specimens of all species are deposited in the collection of the State Museum of Natural History, National Academy of Sciences of Ukraine, Lviv.
Below are the places where the samples were collected in Zakarpattia Region (Fig. 1
Distr ib utio n.Central-eastern Europe (Subias 2018).Ukraine (first record).Currently, the position in Zakarpattia is the easternmost place of the occurrence.Distr ib utio n.Central and southern Europe (Subias, 2018).First record in Ukraine.Presently, Zakarpattia is the easternmost region of its known occurrence.

DISCUSSION
Oribatid species registered for the first time in Ukraine were found in floodplain (A.cf.quadridentata, O. europaea), and in hydromeliorated (O.europaea, A. cf.quadridentata), hygrophilic (O.hygrophila, A. cf.quadridentata) and mesophilic (C.cf.psammophilus) grasslands of the Transcarpathian lowland.These species have been found in Ukraine in similar habitats in which they occur in central Europe (Weigmann & Miko 2006).Only O. europaea was noted in central Europe also in arable field with alluvial soils and in forests (Weigmann & Miko 2006, Ľuptáčik et al. 2012).Finding of the species on the investigated area was expected as they were registered at the neighboring territories (in particular the central European countries).It is also likely that these species will be found in other regions of Ukraine in appropriate biotopes.