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The relationships of the families Colydidae, Monommatidae and Zopheridae are examined. Phylogenetic analyses were carried out using Hennig86 and based on 37 exemplar genera from each of these families and several possible outgroups coded for 59 morphological features (42 adult and 17 larval). Results of the analyses suggest that the three families form amonophilethic group, which may be divided into two subfamilies, Colydiinae and Zopherinae, andthat the latter may be further divided into six tribes: Latometini (trib. nov.), Usechini, Monommatini, Phellopsini (trib. nov.), Pycnomerini (sens. nov.) and Zopherini (sens. nov.). The genera comprising the Zopherinae (excluding Monommatini) are keyed aout and redescribed, and general remarks are made on the biology of the family. Three new genera are described:Orthocerodes gen. nov. (type species: Sarrotrium australis Blackburn, 1891), Australia; Notorhocerus gen. nov.(type species N. valdivianus sp. nov.), Chile and Zopher (type species: Z. ivieisp. nov.), Malaysia. Type species are designated for the genera Docalis Pascoe, 1860 (D.degener Pascoe, 1860) and Elascus (E. crassicornis Pascoe, 1860). Pycnomorphus Matschulski,1858, and Dechomus Jacquelin du Val, 1859, are synonymized with Pycnomeru sErichson, 1842 (syn. nov.). Cotulades brindabellae sp. nov. is described, as is its larva. The following are also proposed: Cotulades okeinom. nov.(=C. pilosus Oke, 1932, not Docalis pilosus Carter, 1928)and C. pilosus (Carter in Wilson, 1928) (Docalis) comb. nov.